noun The fourth stage of prophase of meiosis, during which homologous chromosome pairs begin to separate and chiasmata become visible
noun ↑ "Recherches sur l'Ovogenèse et l'Organogenèse de l'ovaire des Mammifères (Lapin et Homme)", Hans von De Winiwarter, 1900, Archives de biologie; Vol 17
noun ↑ "Studies on chromosomes. VIII. Observations on the maturation‐phenomena in certain Hemiptera and other forms, with considerations on synapsis and reduction", Edmund B. Wilson, 5 October 1912, https://doi.org/10.1002/jez.1400130302